My Love Story with Yamada-kun at Lv999

College student Akane Kinoshita struggles with a wounded heart and the memories her ex left behind after breaking up with her for another girl. She decides to avenge her ex-boyfriend through an in-person event for the MMO Forest of Savior, which they used to play together. While doing so, she encounters Akito Yamada, a gaming legend who also happens to be a guildmate. Akane, who is in need of assistance, convinces the reclusive Yamada to participate in her plan and offer her a shoulder to weep on. Spending time together reveals Akane and Yamada's differences, yet they can't help but notice each other's positive traits on the inside. Their tentative friendship may rise up in a way neither expects as they spend more time together both in and outside of the game.